USC Catalogue 2018-2019 [ARCHIVED CATALOGUE]
Biological Sciences
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Return to: USC Dornsife College of Letters, Arts and Sciences
The Department of Biological Sciences has research faculty with specialties in four disciplines: human and evolutionary biology, marine environmental biology, molecular and computational biology, and neurobiology. A diversity of upper-division undergraduate and graduate courses permits biology majors to choose an emphasis in any of these four disciplines. (See the Neuroscience section of the catalogue for descriptions of the graduate degree in Neuroscience). The department offers both BA and BS degrees in Biological Sciences and an honors program in which a student can earn either a BA or a BS degree in Biological Sciences with Honors. The department also offers both BA and BS degrees in Human Biology and a BS in Quantitative Biology. The BS in Biochemistry is offered as a joint program with the Department of Chemistry. The honors program is available to students who maintain a GPA of 3.5 in the sciences and who have completed their freshman year. The honors program includes research opportunities, seminars and thesis preparation courses. Applications for the Honors Program are available in Allan Hancock Foundation (AHF), Room 105A. Undergraduates in biological sciences have the opportunity to become involved in laboratory or field research by taking research courses for some of their elective units. Minors are offered in biotechnology, computational biology and bioinformatics, human movement sciences, marine biology, and natural science.
At the graduate level, the department offers challenging degree programs that lead to a PhD in Integrative and Evolutionary Biology; Marine Biology and Biological Cceanography; Molecular Biology; and Computational Biology and Bioinformatics. The department also offers progressive MS degrees in Marine and Environmental Biology and Molecular Genetics and Biochemistry.
Allan Hancock Foundation Building 107
(213) 740-2777
FAX: (213) 740-8123
Email (undergraduate programs):
biodept@dornsife.usc.edu
Email (graduate programs):
marinebi@dornsife.usc.edu
molecule@dornsife.usc.edu
ieb@dornsife.usc.edu
Website: dornsife.usc.edu/bisc
Chair: Douglas G. Capone, PhD
Faculty
University Professor and ARCO/William F. Kieschnick Chair in the Neurobiology of Aging and Professor of Gerontology, Biological Sciences, Anthropology and Psychology: Caleb E. Finch, PhD
University Provost Professor of Neurology, Biomedical Engineering and Biological Sciences: Steve Kay, PhD (Neurology)
University Professor and Milo Don and Lucille Appleman Professor of Biological Sciences and Professor of Biological Sciences and Psychology: Larry W. Swanson, PhD
University Professor and USC Associates Chair in Natural Sciences and Professor of Biological Sciences, Computer Science and Mathematics: Michael S. Waterman, PhD
Distinguished Professor of Biological Sciences, Molecular Biology and Biochemistry, and Ester Dornsife Chair in Biological Sciences: Norman Arnheim*, PhD
Distinguished Professor of Chemistry, Biological Sciences, Biochemistry, and Chemical Engineering & Materials Science and Dana and David Dornsife Chair in Chemistry: Arieh Warshel, PhD (Chemistry)
William and Julie Wrigley Chair in Environmental Studies and Professor of Biological Sciences: Douglas G. Capone, PhD
USC Associates Captain Hancock Chair in Marine Science and Professor of Biological Sciences: David A. Caron*, PhD
James E. Birren Chair in Gerontology and Professor of Gerontology and Biological Sciences: Kelvin Davies, PhD (Gerontology)
Elizabeth Garrett Chair in Convergent Bioscience and Provost Professor of Biological Sciences, Biomedical Engineering, Physiology and Biophysics, Stem Cell Biology and Regenerative Medicine, Pediatrics, Radiology and Ophthalmology: Scott Fraser, PhD
McCulloch-Crosby Chair in Marine Biology and Professor of Biological Sciences: Jed A. Fuhrman*, PhD
Wrigley Chair in Environmental Studies and Professor of Earth Sciences and Biological Sciences: Kenneth Nealson, PhD (Earth Sciences)
Robert D. Beyer (‘81) Early Career Chair in Natural Sciences and Associate Professor of Physics, Biological Sciences and Chemistry: Moh El-Naggar*, PhD (Physics and Astronomy)
W. M. Keck Provost Professor of Stem Cell Biology and Regenerative Medicine and Biological Sciences: Andrew McMahon, PhD (Stem Cell Biology and Regenerative Medicine)
Provost Professor of Biological Sciences, Chemistry, Neurology, Physiology & Biophysics, and Chemical Engineering & Materials Science: Ray Stevens, PhD
Provost Professor of Ophthalmology, Neurology, Psychiatry and Behavior Sciences, Radiology, Biomedical Engineering and Biological Sciences: Arthur Toga, PhD
Gabilan Distinguished Professor in Science and Engineering and Professor of Biological Sciences: Susan Forsburg, PhD
Paxson H. Offield Professor of Fisheries Ecology and Professor of Biological Sciences: Dennis Hedgecock, PhD
Gabilan Distinguished Professorship in Science and Engineering and Professor of Biological Sciences: Judith Hirsch, PhD
Gabilan Assistant Professor of Biological Sciences: Carly Kenkel, PhD
Gabilan Assistant Professor of Biological Sciences and Earth Sciences: Naomi Levine, PhD
Gabilan Distinguished Professorship in Science and Engineering and Professor of Biological Sciences and Biomedical Engineering: Jill McNitt-Gray, PhD
Gabilan Assistant Professor of Biological Sciences: Lindsey Schier, PhD
Gabilan Distinguished Professorship in Science and Engineering and Professor of Bioligical Sciences: Lorraine Turcotte, PhD
Professors: Frank Alber, PhD; Jan Amend, PhD (Earth Sciences); Oscar M. Aparicio, PhD; Donald Arnold, PhD; Christopher Boehm, PhD; David Bottjer*, PhD (Earth Sciences); Sarah Bottjer, PhD; Lin Chen, PhD; Xiaojiang Chen, PhD; Vadim Cherezov, PhD (Chemistry); Irene Chiolo, PhD; Pinchas Cohen, PhD (Gerontology); Casey Donovan, PhD; Suzanne Edmands, PhD; Steven Finkel, PhD; Henryk Flashner, PhD (Aerospace and Mechanical Engineering); Myron F. Goodman, PhD; Zach Hall, PhD (Cell and Neurobiology); Albert A. Herrera*, PhD; David Hutchins, PhD; Dale Kiefer, PhD; Chien-Ping Ko, PhD; Peter Kuhn, PhD; Michael Lieber, PhD (Pathology); Emily R. Liman, PhD; Valter D. Longo, PhD (Gerontology); Chi H. Mak*, PhD (Chemistry); Donal T. Manahan, PhD; James W. Moffett, PhD; Sergey Nuzhdin, PhD; John A. Petruska, PhD; Carolyn Phillips, PhD; Michael Quick, PhD; Remo Rohs, PhD; Sergio Sañudo-Wilhelmy, PhD; Richard Simerly, PhD (Medicine); Craig Stanford, PhD; Cornelius W. Sullivan, PhD; Fengzhu Sun, PhD; John Tower, PhD; Alan Watts, PhD
Associate Professors: Liang Chen, PhD; Sean Curran, PhD (Gerontology); Matthew Dean, PhD; Rosa Di Felice, PhD (Physics); Ian Ehrenreich, PhD; Robert Girandola, PhD; Andrew Gracey, PhD; Joseph G. Hacia (Biochemistry and Molecular Mecidine); John F. Heidelberg, PhD; Yan Liu, PhD (Computer Science); David D. McKemy*, PhD; Matthew Michael, PhD; Matthew Pratt, PhD (Chemistry); Andrew Smith, PhD; Eric A. Webb, PhD; Chao Zhang, PhD (Chemistry); Wiebke Ziebis, PhD
Assistant Professors: James Boedicker, PhD (Physics); Mark Chaisson, PhD; Dion Dickman, PhD; Stacey Finley, PhD (Biomedical Engineering); Christoph Haselwandter, PhD (Physics); Bruce Herring, PhD; Samuel Andrew Hires, PhD; Scott Kanoski, PhD; Vsevolod Katritch, PhD; Fabien Pinaud, PhD
Professor (Teaching): Karla B. Heidelberg, PhD
Professor (Research): James Hicks, PhD
Associate Professor (Teaching): Kurt Kwast, PhD
Associate Professors (Research): Linda Duguay, PhD; Feixue Fu, PhD; Le Trinh, PhD
Assistant Professors (Teaching): Christa Bancroft, PhD; Oliver Rizk, PhD; Bruce Yazejian, PhD
Assistant Professors (Research): Peter Calabrese, PhD; Phuong Pham, PhD
Master Lecturer: Gudrun Floyd, MS
Lecturers: Stephanie Bogart, PhD; Nancy Castro, PhD; Tamara Espinet, MS; Raffaella Ghittoni, PhD; Michael Hadjidaniel, PhD; Grayson Jaggers, PhD; Refael Levi, PhD; Gioia Polidori Francisco, PhD
Emeritus Professors: Michael Appleman, PhD; Robert Baker, PhD; John Callaghan, PhD; Richard Deonier, PhD; William O. McClure*, PhD; Russel Zimmer, PhD
*Recipient of university-wide or college teaching award.
Undergraduate Degrees
Advisement
Advisement in the Department of Biological Sciences is required each semester. First semester freshman and transfer advisement takes place during orientation. Advisement in all remaining semesters takes place during the pre-registration period. The undergraduate coordinator forwards advisement appointment information each semester to all students in biological sciences and biochemistry.
Pre-Medical and Other Pre-Professional Preparation
The department offers specially planned courses within the biological sciences to prepare students for admission to professional schools (medicine, dentistry, veterinary science, occupational therapy, physical therapy, pharmacy, optometry, public health), paramedical sciences (medical technology, physician’s assistant, clinical and public health microbiology, clinical biochemistry), naturalist and environmental positions in the public and private sectors, jobs in industry (biotechnology), and graduate study (basic biological and biomedical fields). With the proper selection of courses under the guidance of the Department of Biological Sciences and the USC Rossier School of Education, the BS degree satisfies the California requirements for secondary school teaching in the life sciences.
Honor Society
The Department of Biological Sciences offers membership in Phi Sigma, a national honor society, to selected biology majors (Alpha Alpha Chapter at USC.) Phi Sigma is devoted to the promotion of research and academic excellence in the biological sciences. Students with a GPA above 3.0 who have interest in research and have completed core requirements for the first two years in biological sciences are eligible. Major activities range from presentation of papers by members and lectures by outside speakers to field trips, laboratory demonstrations and joint research projects.
Maymester Courses on Catalina Island
The Biological Sciences Department in conjunction with the USC Wrigley Institute for Environmental Studies (WIES) sponsors 4-week Maymester programs at USC’s Phillip K. Wrigley Marine Sciences Center (WMSC) on Santa Catalina Island. The Maymester courses focus on conservation biology, animal physiology, aquatic microbiology and field techniques in marine biology and biological oceanography. These courses are open to all biology majors as well as students in other departments and other institutions with a strong biology background. Students are primarily in their junior or senior years and may participate in any of the courses offered.
All the courses are taught by USC faculty and supported by USC graduate student teaching assistants. The classes are specialized to take advantage of the unique facilities and setting of Santa Catalina Island.
Students live on Catalina Island for the entire length of the course. Rates for room and board at the USC Wrigley Marine Science Center are comparable to those on campus. For those wishing to return to the mainland on the weekend — free transportation is provided each Friday to leave and each Monday to return.
For more information, students should contact the instructor(s) for the course of interest. Additional information can also be found at dornsife.usc.edu/wrigley/education/.
Minor in Craniofacial and Dental Technology
For a complete listing of course requirements, see the Herman Ostrow School of Dentistry of USC .
Minor in Biotechnology
The Marshall School and the departments of biological sciences and chemistry in the USC Dornsife College of Letters, Arts and Sciences jointly offer the minor in biotechnology. This minor brings essential knowledge in the basic sciences together with the corporate skills needed in a rapidly growing industry. The minor is especially well suited for the business, biology, chemistry or engineering student seeking a career in business and/or the biomedical/bio-technical sciences. See Biotechnology Minor for a list of required courses.
Graduate Degrees
Degree Programs in Biological Sciences
The graduate programs in biology provide education and training of biologists interested in living systems ranging from cellular to ecosystem levels of organization, investigated by laboratory or fieldwork. Courses and faculty research interests allow a multidisciplinary approach. A number of additional research areas are provided by adjunct faculty from other institutions, including the Los Angeles County Museum of Natural History and Children’s Hospital Los Angeles. Students develop the ability to formulate and test hypotheses, integrating information and concepts in the completion of a dissertation (PhD). A qualifying exam committee is formed for each student during the first year to develop a particular program of course work and research, and to evaluate the student’s progress. Specific information about the options in biological sciences can be obtained by requesting information brochures or online at dornsife.usc.edu/bisc.
Admission Requirements
Applicants must have a bachelor’s degree in a natural science (preferably biology) from an accredited four year college or university, or in mathematics or engineering; required background courses include organic chemistry, general physics and mathematics through integral calculus. Applicants are evaluated by their transcripts and GPA; scores on the GRE General Test; three letters of recommendation; and a statement of interest. A faculty member must serve as initial sponsor and adviser for admission to marine biology and biological oceanography (MBBO) and integrative and evolutionary biology (IEB). Applicants who are accepted but judged to have minor deficiencies are expected to correct them within the first year.
Degree Requirements
These degrees are awarded under the jurisdiction of the Graduate School. Refer to the Requirements for Graduation section and The Graduate School section of the catalogue for general regulations. All courses applied toward the degrees must be courses accepted by the Graduate School.
Molecular and Computational Biology
This program is designed to train the participants intensively in the concepts and experimental methodologies of molecular biology and biochemistry. The subject matter is organized in an integrated fashion (lectures, seminars and laboratory) to present fundamental information on the biochemistry, biophysics, genetics and development of cells from a variety of different organisms. Primary emphasis is on the relationship between structure and function at different integrative and functional levels. The program offers a PhD in Molecular Biology and a PhD in Computational Biology and Bioinformatics. Molecular and computational biology (MCB) students are required to complete at least two laboratory rotations in their first year. Applications may be accessed online at dornsife.usc.edu/bisc/mcb/.
Admission Requirements
Applicants are expected to have a bachelor’s degree or equivalent in a cognate area such as biology, chemistry, physics, engineering, bacteriology, computer science, or bioinformatics. Undergraduate work should include a basic course in biology, basic physics, physical chemistry, organic chemistry, biochemistry and calculus. Students who are deficient in any of these may be required to correct the deficiency during the first two years of graduate study. Courses taken to correct these deficiencies are usually not credited toward the degree. The student must submit letters of recommendation from at least three faculty members who can evaluate the promise of the student for graduate work and independent research. The applicant must take the GRE General Test prior to acceptance.
Degree Requirements
These degrees are awarded under the jurisdiction of the Graduate School. Refer to the Requirements for Graduation section and The Graduate School section of this catalogue for general regulations. All courses applied toward the degrees must be courses accepted by the Graduate School.
Bachelor’s Degree
Minor
Master’s Degree
Doctoral Degree
Biological Sciences
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BISC 101Lgx Cellular and Molecular Biology Units: 4
Terms Offered: FaSp
Cellular and molecular biology with examples related to human biology and diseases. Fundamental life processes examined at the genetic, cellular and molecular levels.
Satisfies New General Education in Category D: Life Sciences
Satisfies Old General Education in Category III: Scientific Inquiry
Credit Restriction: Not for major credit
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 102Lgx Humans and Their Environment Units: 4
Terms Offered: FaSp
An examination of the physical and biological laws that influence agriculture, pollution, population dynamics (including humans), climate, biodiversity and ecosystem structure and function.
Satisfies New General Education in Category D: Life Sciences
Satisfies Old General Education in Category IV: Science and Its Significance
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 103Lgx General Biology for the Environment and Life Units: 4
Study of common skills in biology, including basics of evolution, systematics, ecology, genetics, biochemistry and molecular biology, physiology, and anatomy.
Satisfies New General Education in Category D: Life Sciences
Instruction Mode: Lecture, Lab
Grading Option: Letter
-
BISC 104Lgx How the Body Works Units: 4
Terms Offered: Fa
Structure and function of the human body, including the role of organ systems, tissues, and cells in normal function. Malfunctions relating to disease, substance abuse and lifestyle.
Satisfies New General Education in Category D: Life Sciences
Satisfies Old General Education in Category III: Scientific Inquiry
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 108L Special Laboratory I Units: 1
Laboratory component for BISC 120 for entering freshmen or transfer students with advanced placement or equivalent lecture credit from another institution.
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 109L Special Laboratory II Units: 1
Laboratory component for BISC 220 for entering freshmen or transfer students with advanced placement or equivalent lecture credit from another institution.
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 110g Good Genes, Bad Genes Units: 4
Terms Offered: Sp
Introduction to the scientific method; basic instruction in molecular biology with emphasis on how biological research advancement has impacted medicine, commerce and society in general.
Recommended Preparation: High school biology
Satisfies New General Education in Category D: Life Sciences
Instruction Mode: Lecture, Discussion
Grading Option: Letter
-
BISC 115Lxg The Biology of Food Units: 4
Terms Offered: FaSp
Exploration of molecular biology, biochemistry, microbiology, nutrition and the history of biology through the study of food, focusing on food’s relationship to the biological world.
Satisfies New General Education in Category D: Life Sciences
Credit Restriction: Not available for credit for Biology majors
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 120Lg General Biology: Organismal Biology and Evolution Units: 4
Terms Offered: Fa
In-depth survey of key topics related to advances in our knowledge of the diversity of life and evolution; origin of life; eukaryotes/prokaryotes; ecology.
Satisfies New General Education in Category D: Life Sciences
Satisfies Old General Education in Category III: Scientific Inquiry
Duplicates Credit in BISC 112L, BISC 113L, and BISC 121L.
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 121Lg Advanced General Biology: Organismal Biology and Evolution Units: 4
Terms Offered: Fa
Equivalent to BISC 120 , but taught at a higher level for exceptionally well-prepared students. Admission to the course by departmental approval only.
Corequisite: CHEM 115aLg .
Satisfies New General Education in Category D: Life Sciences
Satisfies Old General Education in Category III: Scientific Inquiry
Duplicates Credit in BISC 112L, BISC 113L, and BISC 120L.
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 140g Our Blue Planet in a Changing Climate Units: 4
Terms Offered: Sp
Introduction to biological oceanography and marine ecosystems. Impact of climate changes on marine ecosystems. Analysis of ocean-related case studies being discussed by mainstream media.
Satisfies New General Education in D Life Sciences
Instruction Mode: Lecture, Discussion
Grading Option: Letter
-
BISC 150Lgx The Nature of Human Health and Disease Units: 4
Terms Offered: FaSp
The human organism; the nature of inherited and acquired diseases; the biological and societal basis for the AIDS epidemic; therapy, drug design and the future.
Satisfies New General Education in Category D: Life Sciences
Satisfies Old General Education in Category IV: Science and Its Significance
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 180Lgx Evolution Units: 4
Terms Offered: Sp
Changes in the physical and biological universe over time; origins of life, dinosaurs, human evolution. Implications of evolutionary mechanisms and mass extinctions for human survival.
Satisfies New General Education in Category D: Life Sciences
Satisfies Old General Education in Category IV: Science and Its Significance
Credit Restriction: Not available for credit in any major.
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 193 Introduction to Research I Units: 1
Terms Offered: Fa
A series of lectures and discussions at which faculty of the department introduce their research activities to students entering biology and related majors.
Corequisite: BISC 120Lg or BISC 121Lg
Instruction Mode: Lecture
Grading Option: Credit/No Credit
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BISC 194 Introduction to Research II Units: 1
Terms Offered: Sp
A series of lectures and discussions at which faculty of the department introduce their research activities to students entering biology and related majors.
Corequisite: BISC 220Lg or BISC 221Lg
Instruction Mode: Lecture
Grading Option: Credit/No Credit
-
BISC 199 Neuroscience Colloquium Units: 1
Terms Offered: Sp
Introduction to research activities conducted by Neuroscience faculty at USC.
Instruction Mode: Lecture
Grading Option: Credit/No Credit
Crosslisted as NEUR 199 -
BISC 220Lg General Biology: Cell Biology and Physiology Units: 4
Terms Offered: Sp
In-depth survey of key topics related to advances in our knowledge of cellular biology and physiology; cell composition/metabolism; gene action; organism structure and function.
Recommended Preparation: high school chemistry; BISC 120Lg or BISC 121Lg .
Satisfies New General Education in Category D: Life Sciences
Duplicates Credit in BISC 110L, BISC 111L, and BISC 221L.
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 221Lg Advanced General Biology: Cell Biology and Physiology Units: 4
Terms Offered: Sp
Equivalent to 220L, but taught at a higher level for exceptionally well-prepared students. Admission to the course by departmental approval only.
Prerequisite: BISC 120Lg or BISC 121Lg ;
Corequisite: CHEM 105bL or CHEM 115bL .
Satisfies New General Education in Category D: Life Sciences
Duplicates Credit in BISC 110L, BISC 111L, and BISC 220L.
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 230Lgx The Biology of the Brain Units: 4
Terms Offered: FaSpSm
The structure and function of the mammalian brain including the role of the brain in regulating behavior, both in normal and diseased states; in relation to mind; and in comparison with machine forms of intelligence.
Satisfies New General Education in Category D: Life Sciences
Satisfies Old General Education in Category IV: Science and Its Significance
Credit Restriction: Not available for major credit
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 290 Introduction to Biological Research Units: 2 or 4
Max Units: 4.0
Terms Offered: FaSpSm
Experience in basic techniques through supervised research in the research laboratory of a departmental faculty member.
Prerequisite: BISC 120Lg or BISC 121Lg ; BISC 220Lg or BISC 221Lg ; CHEM 105bL or CHEM 115bL ; departmental approval.
Instruction Mode: Lecture
Grading Option: Credit/No Credit
-
BISC 300L Introduction to Microbiology Units: 4
Terms Offered: Sp
Comparative approach to bacteria, Archaea and viruses; their structure, life cycles, geochemical activity, ecology and nutrition. Fundamentals of metabolism and microbial genetics.
Prerequisite: BISC 320 , CHEM 322aL or CHEM 325aL .
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 305 Introduction to Statistics for Biologists Units: 4
Terms Offered: Fa
Statistical methods in biological science and medicine, including populations and samples, random sampling, confidence intervals, paired samples and regression.
Instruction Mode: Lecture, Discussion
Grading Option: Letter
-
BISC 307L General Physiology Units: 4
Terms Offered: Sp
Physiological functions of the circulatory, digestive, endocrine, integumentary, musculoskeletal, nervous, respiratory, and urogenital systems of animals.
Prerequisite: BISC 220Lg or BISC 221Lg .
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 312x Molecular Biochemistry Units: 4
Terms Offered: FaSp
Structure and function of biological macromolecules; major metabolic pathways including glycolysis and photosynthesis. DNA replication, repair, and recombination; gene expression, regulation and epigenetics.
Recommended Preparation: BISC 220 or BISC 221
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 313L Evolution and Population Genetics Units: 4
Terms Offered: Sp
History of evolutionary thought; molecular basis for evolution; dynamics of genes in populations; speciation and macro evolution; patterns of evolution.
Prerequisite: BISC 220Lg or BISC 221Lg ; BISC 120Lg or BISC 121Lg ;
Recommended Preparation: BISC 320L , BISC 325 , and familiarity with algebra, basic chemistry, and basic physics.
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 315L Introduction to Ecology Units: 4
Terms Offered: Fa
Organism-environment interactions; dynamics of populations, communities, and ecosystems; evolutionary forces.
Prerequisite: BISC 120Lg or BISC 121Lg .
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 320L Molecular Biology Units: 4
Terms Offered: Fa
Structure and synthesis of nucleic acids and proteins; molecular biology of prokaryotes and eukaryotes; principles of genetics and cell biology.
Prerequisite: CHEM 105bL or CHEM 115bL .
Satisfies New General Education in D Life Sciences
Duplicates Credit in BISC 311.
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 321 Multidisciplinary Seminar: Science, Technology and Society Units: 2
Terms Offered: Sp
Builds upon a basic science background to provide students with an awareness of cutting edge scientific research, its technological applications and its societal ramifications.
Prerequisite: (BISC 120Lg or BISC 121Lg or BISC 220Lg or BISC 221Lg ) and (CHEM 105aLg or CHEM 115aLg ) and (PHYS 135ag or PHYS 151Lg )
Duplicates Credit in former MDA 321
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 325 Genetics Units: 4
Terms Offered: Fa
Transmission genetics and genotype/phenotype; mapping methods; complex traits; genetics of human disease and population genetics.
Prerequisite: BISC 120Lg or BISC 121Lg ; BISC 220Lg or BISC 221Lg ; BISC 320L ; CHEM 322aL ;
Corequisite: CHEM 322bL or CHEM 325bL .
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 330L Biochemistry Units: 4
Terms Offered: FaSp
Basic biochemical principles; classes of molecules - structure and function; cellular energetics.
Prerequisite: CHEM 322aL
Duplicates Credit in former BISC 316
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
Crosslisted as CHEM 330 -
BISC 335 Science, Health and the Environment Units: 4
Terms Offered: Sp
(Enroll in ENST 335 )
-
BISC 352 Conservation Biology Units: 4
Terms Offered: Sp
Principles of conservation science in marine and terrestrial ecosystems with emphasis on protecting biological diversity and balancing the needs of nature with those of humans.
Recommended Preparation: Introductory course in biology such as BISC 103 , BISC 120 or BISC 121 , or AP Biology credit.
Instruction Mode: Lecture, Lab
Grading Option: Letter
Crosslisted as ENST-352 -
BISC 369L Ecology and the Natural History of California Units: 4
Terms Offered: Sp
Marine, freshwater, and terrestrial communities of California. Life histories, morphology, special evolutionary adaptations. Relationships between organisms and their biological-physical-chemical environment. Offered on Catalina. Emphasis on field biology.
Prerequisite: BISC 120Lg or BISC 121Lg .
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 371L Molecular Approaches to the Diversity of Life Units: 4
Patterns of evolutionary change investigating the molecular basis of heredity utilizing DNA data. History, principles and application of molecular systematics, and genetic variation. Taught on Catalina Island.
Prerequisite: BISC 120Lg or BISC 121Lg ; BISC 220Lg or BISC 221Lg ;
Recommended Preparation: BISC 320L .
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 390 Special Problems Units: 1, 2, 3, 4
Supervised, individual studies. No more than one registration permitted. Enrollment by petition only.
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 403 Advanced Molecular Biology Units: 4
Terms Offered: Fa
Molecular mechanisms and control of DNA replication, DNA repair, recombination, gene expression, cell growth, and development in prokaryotic and eukaryotic organisms, from bacteria to humans.
Prerequisite: BISC 320
Recommended Preparation: BISC 313 or BISC 325 .
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 405L General Embryology Units: 4
Terms Offered: Fa
Vertebrate and human development: cellular differentiation; germ cell development and growth; hormonal regulation of reproductive cycles; cleavage through neurulation and subsequent development of primary organs.
Prerequisite: BISC 120Lg or BISC 121Lg ; BISC 220Lg or BISC 221Lg ;
Recommended Preparation: two from BISC 313 , BISC 320L , BISC 325 and BISC 330L .
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 406L Biotechnology Units: 4
Terms Offered: Fa
Techniques in molecular biology and biochemistry applied to prokaryotic and eukaryotic model systems; applications of recombinant DNA and genomic technology.
Prerequisite: BISC 320L ;
Recommended Preparation: BISC 313 or BISC 325 .
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 407 Cellular and Molecular Neuroscience Units: 4
Terms Offered: FaSp
An examination of individual nerve cell development, target location, and establishment of functional synapses; how dysfunction in these processes contributes to neurological and neuropsychiatric diseases.
Recommended Preparation: BISC 421
Instruction Mode: Lecture, Discussion
Grading Option: Letter
Crosslisted as NEUR 407 -
BISC 408 Systems Neuroscience: From Synapses to Perception Units: 4
Terms Offered: Sp
Sensory systems to illustrate basic concepts regarding the functional organization of the brain, from the microscopic arrangement of neural circuits to global processes such as perception.
Prerequisite: BISC 421 .
Instruction Mode: Lecture, Discussion
Grading Option: Letter
Crosslisted as NEUR-408 -
BISC 410 Applications of Molecular Biology to Medicine Units: 4
Terms Offered: Sp
Advances and trends in the understanding, diagnosis and treatment of human diseases.
Prerequisite: BISC 330L .
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 411 Advanced Cell Biology Units: 4
Terms Offered: FaSpSm
The synthesis, transport and assembly of the complex structures that mediate eukaryotic cellular function. Electrical and biochemical mechanisms underlying intercellular communication.
Prerequisite: BISC 220Lg or BISC 221Lg ; BISC 320L .
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 414 Biology of Cancer Units: 4
Terms Offered: Sp
Focus on the advances in molecular biology of cancer, from fundamental molecular signaling pathways to DNA repair to stem cell biology, through primary research literature reviews.
Prerequisite: BISC 325 .
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 419L Environmental Microbiology Units: 4
Terms Offered: FaSpSm
Qualitative and quantitative appraisal of microbial activities in pure and contaminated environments; microbial community and its development; interspecific relationships; effects of microorganisms on their surroundings.
Prerequisite: BISC 330L ;
Recommended Preparation: BISC 300L .
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 421 Neurobiology Units: 4
Terms Offered: Fa
Structure, function, and development of nervous systems; neural integration and mechanisms of behavior; organization and operation of brains.
Prerequisite: BISC 220Lg or BISC 221Lg
Instruction Mode: Lecture, Discussion
Grading Option: Letter
Crosslisted as NEUR 421 -
BISC 422L Neurobiology Laboratory Units: 2
Terms Offered: Sp
Experimentation on excitable cells, synapses, and neural circuits; intracellular and extra cellular techniques for recording, stimulation, and identification of nerve and muscle cells.
Corequisite: BISC 421 .
Instruction Mode: Lab Required
Grading Option: Letter
-
BISC 423 Epilepsy to Ecstasy: Biological Basis of Neurological Disorders Units: 4
Terms Offered: Sp
Examination of various neurological disorders originating from developmental signaling and/or anatomical abnormalities.
Prerequisite: BISC 421
Instruction Mode: Lecture
Grading Option: Letter
Crosslisted as NEUR 423 -
BISC 424 Brain Architecture Units: 4
Terms Offered: Sp
How the parts of the brain are interconnected to form a complex biological computer, from historical, evolutionary, and developmental perspectives.
Prerequisite: BISC 421
Instruction Mode: Lecture
Grading Option: Letter
Crosslisted as NEUR 424 -
BISC 425 Advanced Genetics through the Primary Scientific Literature Units: 4
Terms Offered: Sp
Literature-based seminar in current and classical topics in genetics.
Prerequisite: BISC 325 .
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 426 Principles of Neural Development Units: 4
Terms Offered: Sp
Basic phenomena and principles of neural development, their relation to functional development of neural circuits, behavior, and disease. General concepts and experimental approaches are emphasized.
Prerequisite: BISC 421 .
Instruction Mode: Lecture
Grading Option: Letter
Crosslisted as NEUR-426 -
BISC 427 The Global Environment Units: 4
Terms Offered: FaSpSm
Earth’s development as a habitable planet, from origin to human impacts on global biogeochemical cycles in the ocean, land, atmosphere. Discussion of environmental alternatives.
Prerequisite: (BISC 103 or BISC 120 or BISC 121 and (CHEM 103 or CHEM 105b or CHEM 115b )
Registration Restriction: Open only to Biological Sciences, Environmental Studies, Earth Sciences, and Geological Sciences majors
Instruction Mode: Lecture
Grading Option: Letter
Crosslisted as ENST 427, GEOL 427 -
BISC 428 The Biology of Tropical Diseases Units: 4
Max Units: 8.0
Terms Offered: Sm
Biological and biochemical approaches to the prevention and treatment of infectious and chronic tropical diseases. Course is offered off campus and summer only.
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 429 Cancer Immunology Units: 4
Terms Offered: Sp
In-depth study of the role of the immune system in oncogenesis. Discussion of cellular transformation, immune surveillance, immune-based therapies, and new approaches in cancer treatment.
Prerequisite: BISC 220Lg or BISC 221Lg
Recommended Preparation: Familiarity with Molecular Biology, Genetics, Principles of Immunology
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 431L Aquatic Microbiology Units: 4
Terms Offered: FaSpSm
Introduction to the habitat, phylogenetic, physiological and metabolic diversity of microbial life in aquatic environments.
Prerequisite: BISC 220Lg or BISC 221Lg
Duplicates Credit in BISC 419L
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 434 Introduction to Genome Science Units: 4
Terms Offered: Sp
Characterization of the sequence, function and evolution of genomes; study of the molecular basis of phenotypes and the relationship between genomics and synthetic biology.
Recommended Preparation: BISC 325
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 435 Advanced Biochemistry Units: 4
Terms Offered: Sp
Macro molecular structure and function; enzymology; metabolic regulation.
Prerequisite: BISC 330L .
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 437L Comparative Physiology of Animals Units: 4
Terms Offered: Sp
Control of the internal environment of animals in relation to their external environment. Thermal regulation, osmoregulation, excretion, and ion balance. Offered on Catalina.
Prerequisite: BISC 120 or BISC 121 ; BISC 220 or BISC 221 ;
Recommended Preparation: two from BISC 313 , BISC 320L , BISC 325 and BISC 330L .
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 438 Nutritional Biochemistry Units: 4
Terms Offered: FaSp
Metabolism of carbohydrates, proteins, fats. Biochemical basis of nutrition’s impact on metabolic functions, including vitamins, minerals and biologically active non-nutrients; medical aspects of nutrition.
Prerequisite: BISC 330L
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 440 Biodemography of Aging Units: 4
(Enroll in GERO 440 )
-
BISC 444 Practical Analysis of Biological Data in R Units: 2
Terms Offered: FaSp
Instruction in the open-source statistical program environment R to analyze biological data; manipulation of large datasets and customization of statistical tests using simulations.
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 445L Fundamentals of Vertebrate Biology Units: 4
Terms Offered: Sp
Evolution and comparative anatomy of vertebrates.
Prerequisite: BISC 120Lg or BISC 121Lg ; BISC 220Lg or BISC 221Lg ;
Recommended Preparation: two from BISC 313 , BISC 320L , BISC 325 and BISC 330L .
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 447L Island Biogeography and Field Ecology Units: 4
Terms Offered: Sp
Biogeography, geology, ecology, climate, flora, and fauna of terrestrial and marine environments of Catalina and the Channel Islands including laboratory and field techniques of ecology. Taught on Catalina Island.
Prerequisite: BISC 120Lg or BISC 121Lg .
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 450L Principles of Immunology Units: 4
Terms Offered: Fa
Immune processes, humoral and cellular; immunoglobulins; antibody formation; antigen-antibody interactions; immune dyscrasias; transplantation and tumor immunology; basic hematology and immuno-hematology.
Prerequisite: BISC 220Lg or BISC 221Lg .
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 455L Molecular Approaches to Microbial Diversity — Catalina Semester Units: 4
Terms Offered: Fa
Overview and practical application of genetic and immunological techniques for examining diversity and community structure of natural microbial assemblages in aquatic ecosystems.
Prerequisite: BISC 320L ;
Corequisite: BISC 431L .
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 456L Conservation Genetics Units: 4
Terms Offered: Sp
Biological principles underlying conservation including ecology, evolution, genetics and biogeography. Covers both marine and terrestrial environment, with special emphasis on island biology. Catalina semester only.
Prerequisite: BISC 120Lg or BISC 121Lg ; BISC 220Lg or BISC 221Lg ;
Recommended Preparation: BISC 320L ; BISC 313 or BISC 325 .
Duplicates Credit in BISC 373L.
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
Crosslisted as ENST-456 -
BISC 457L Methods in Marine Biology and Biological Oceanography Units: 4
Terms Offered: Sp
Introduction to standard methods used in oceanography and marine biology through a combination of lectures, laboratory exercises and field experiences.
Prerequisite: BISC 103 or BISC 120 or BISC 121
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 460 Seminar in Marine and Environmental Biology Units: 2
Max Units: 4.0
Terms Offered: FaSp
Topical seminar in marine and environmental biology.
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 461 Seminar in Molecular and Computational Biology Units: 2
Terms Offered: FaSpSm
Topical seminar in molecular and computational biology.
Instruction Mode: Lecture
Grading Option: Letter
Crosslisted as GERO-461 -
BISC 462 Seminar in Neurobiology Units: 2
Max Units: 04
Terms Offered: FaSp
Topical seminar in neurobiology.
Registration Restriction: Open only to juniors, seniors, master and doctoral students
Instruction Mode: Lecture
Grading Option: Letter
Crosslisted as NEUR 462 -
BISC 469L Marine Biology Units: 4
Terms Offered: Sp
Oceanography and marine biology, sampling techniques, evolutionary adaptations, morphology, systematics.
Prerequisite: BISC 103Lgx or BISC 120Lg or BISC 121Lg
Recommended Preparation: BISC 473L
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
-
BISC 473L Biological Oceanography Units: 4
Terms Offered: Fa
Biological, physical, chemical dynamics and analysis of the ocean; primary production of phytoplankton, secondary production by zooplankton, bacterial remineralization; physiology, ecology of fishes, marine mammals.
Prerequisite: BISC 103 or BISC 120 or BISC 121Lg
Registration Restriction: Junior, Senior, or with permission from the instructor
Instruction Mode: Lecture, Lab
Grading Option: Letter
-
BISC 474L Ecosystem Function and Earth Systems Units: 4
Terms Offered: Fa
General principles of ecosystem function, energy flow and materials cycling in marine systems at various scales and the importance of microbial processes in these systems. Taught on Catalina Island.
Prerequisite: BISC 120Lg or BISC 121Lg .
Instruction Mode: Lecture, Lab Required
Grading Option: Letter
Crosslisted as GEOL-474 -
BISC 478 Computational Genome Analysis Units: 4
Terms Offered: Sp
Introduction to and applications of algorithms and statistics to genome analysis. Analysis of physical and genetic maps, DNA sequencing, sequence comparisons, DNA chips.
Instruction Mode: Lecture
Grading Option: Letter
Crosslisted as QBIO 478 -
BISC 480 Developmental Biology Units: 4
Terms Offered: FaSp
Basic mechanisms of animal development are considered at different levels of analysis. Emphasis is on molecular, genetic, and cellular processes underlying vertebrate and invertebrate development. General concepts and evolutionary mechanisms are emphasized.
Prerequisite: BISC 220Lg or BISC 221Lg .
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 481 Structural Bioinformatics: From Atoms to Cells Units: 4
Terms Offered: Fa
Introduction to computational and experimental methods for structural bioinformatics. Analysis of macro molecules and their complexes.
Instruction Mode: Lecture
Grading Option: Letter
Crosslisted as QBIO 481 -
BISC 483 Geobiology and Astrobiology Units: 4
Terms Offered: Sp
Relationships between microbiota and the earth environment including the hydrosphere, lithosphere and atmosphere, with consideration of the potential for life on other planets.
Prerequisite: BISC 120Lg , CHEM 105bL .
Instruction Mode: Lecture
Grading Option: Letter
Crosslisted as GEOL-483 -
BISC 485 Advanced Seminar in Bacterial Survival and Evolution Units: 4
Terms Offered: Sp
Literature-based seminar in current topics in microbial evolution and adaptation.
Prerequisite: BISC 120 or BISC 121 , BISC 220 or BISC 221 , BISC 320 , CHEM 322a or CHEM 325a .
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 486 Regenerative Medicine: Principles, Paradigms and Practice Units: 4
Terms Offered: Fa
Examination of molecular and cellular processes that generate, maintain and repair body systems. Special emphasis on stem cell biology with clinical applications to regenerative medicine.
Prerequisite: (BISC 220Lg or BISC 221Lg ) or BISC 320L
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 490x Directed Research Units: 1, 2, 3, 4, 5, 6, 7, 8
Max Units: 12.0
Terms Offered: FaSpSm
Individual research and readings.
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 493x Honors Seminar Units: 1
Max Units: 4.0
Terms Offered: FaSp
Prerequisite: BISC 120Lg or BISC 121Lg ; BISC 220Lg or BISC 221Lg .
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 494x Honors Thesis Units: 2
Terms Offered: FaSp
Prerequisite: BISC 493 .
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 499 Special Topics Units: 2, 3, 4
Max Units: 8.0
Terms Offered: FaSp
Lecture and discussion in specialized areas of the biological sciences. Students cannot register more than twice for this course.
Recommended Preparation: BISC 220Lg or BISC 221Lg , BISC 320L , BISC 325 and BISC 330L .
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 502a Molecular Genetics and Biochemistry Units: 4
Terms Offered: Fa
Current genetic and biochemical analysis of replication, recombination, mutagenesis, and repair. Fundamentals of transcription and regulation of gene expression. Recent applications of genetic engineering and genome analysis.
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 502b Molecular Genetics and Biochemistry Units: 4
Terms Offered: FaSp
Current genetic and biochemical analysis of replication, recombination, mutagenesis, and repair. Fundamentals of transcription and regulation of gene expression. Recent applications of genetic engineering and genome analysis.
Instruction Mode: Lecture, Discussion
Grading Option: Letter
-
BISC 504L Laboratory Techniques in Cellular and Molecular Biology Units: 1, 2, 3, 4
Max Units: 8.0
Terms Offered: FaSp
Rotation of graduate students through Molecular Biology research laboratories to learn the major technological skills required in the field.
Instruction Mode: Lecture, Lab Required
Grading Option: Credit/No Credit
-
BISC 505 Genomics and Molecular Genetics Units: 4
Terms Offered: Sp
Molecular genetics (mutagenesis, repair, recombination, and gene regulation) from quantitative and mechanistic approaches. Simple and complex genome analysis using recombinant DNA, physical, and computational techniques.
Recommended Preparation: BISC 502b .
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 511 Integrative Biology Units: 4
Terms Offered: Fa
Current topics in integrative biology including form, function and energy use throughout the lifespan in the context of genetics, natural selection and ecology.
Duplicates Credit in the former BISC 510a.
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 512 Evolutionary Biology Units: 4
Terms Offered: Sp
Survey of current topics in evolutionary biology; genetics, natural selection, ecology; emphasis on higher order complex questions of lifespan, form, function, and energy use.
Duplicates Credit in the former BISC 510b.
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 515 Evolution and Human Biology Units: 4
Terms Offered: Fa
Topics in evolution and human biology with emphasis on life span, form, function and energy use in the context of genetics, natural selection and ecology.
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 519 Recent Advances in Neurobiology and Endocrinology of Aging Units: 2, 4
(Enroll in GERO 519 )
-
BISC 520 Recent Advances in Neurobiology Units: 2 or 4
Max Units: 12.0
Terms Offered: Fa
Lectures on selected topics in neurobiology. Registration restricted to three semesters.
Prerequisite: graduate status in departmental program or departmental approval.
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 521 Hearing and Communication Neuroscience Units: 4
Terms Offered: Sp
A basic grounding in broad aspects of the neuroscience of hearing and vocal communication.
Prerequisite: BISC 421 and NSCI 524
Instruction Mode: Lecture
Grading Option: Letter
Crosslisted as NSCI 521 -
BISC 522 Nonlinear Dynamical Systems, Vibrations, and Chaos Units: 3
Terms Offered: Sp
(Enroll in AME 520 )
-
BISC 529 Seminar in Marine Biology Units: 1
Max Units: 4.0
Terms Offered: FaSp
Instruction Mode: Lecture
Grading Option: Credit/No Credit
-
BISC 530 Advanced Seminar in Plankton Biology Units: 2
Terms Offered: FaSp
An overview of phytoplankton and zooplankton taxa, their morphologies and life histories using material collected from the local environment off LA and near the Phillip K. Wrigley Marine Science Center on Catalina Island.
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 531 Advanced Seminar on the Physiology of Marine Organisms Units: 2
Terms Offered: FaSp
Physiological processes dictate survival potential, growth rates, and many other biological processes that affect the distribution of species in the oceans. Emphasis on the diverse environmental factors that influence physiological adaptations of marine organisms. Examples from a wide variety of marine organisms, from bacterial to animals, will be studied.
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 532 Advanced Seminar in Molecular and Microbial Ecology Units: 2
Terms Offered: FaSp
Microorganisms dominate biological processes in the ocean. These species pose significant problems for estimating species diversity, abundance and activity. Examination of modern molecular biological approaches for analyzing aquatic microbial communities and their ecological roles.
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 533 Advanced Seminar in Remote Sensing and Modeling Units: 2
Modern oceanographic methods for making remote measurements of aquatic and terrestrial ecosystems using satellite imagery and other means. Integrating these data into models that describe ecosystem structure and enable interpretation of ecosystem function.
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 534 Advanced Seminar in Population Genetics of Marine Organisms Units: 2
An overview of the theory underlying population and quantitative genetics, with applications to marine systems. Basic evolutionary mechanisms (mutation, migration, drift, selection, nonrandom mating) and modern evidence for their roles in structuring genetic variation within and among marine populations.
Instruction Mode: Lecture
Grading Option: Letter
-
BISC 535 Seminar in Physiology Units: 2
Max Units: 8.0
Terms Offered: FaSp
Instruction Mode: Lecture
Grading Option: Letter
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